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This text represents a synthetic approach to the understanding of the evolution of life history variation using the three types of environment (constant, stochastic, predictable) as the focus under which the theory is developed and tested. The theme of the book is that an understanding of evolutionary change requires analysis at both the genetic and phenotypic levels, and that the environment plays a central role in such analyses. Intended for graduate students and researchers, the book's emphasis is on assumptions and testing of models. Mathematical processes are described, but mathematical derivations are kept to a minimum. Each chapter includes a summary, and boxes provide supplementary material.